From BlenderWiki
Themes
Blender allows the utilization of themes to define custom interface colors and icons. You can manage themes from here, and two are built-in: Default and Rounded. Many others are available from the Internet, such as “Dark Alpha” and “GONX”. Each theme is a Python script, usually ending in *theme.py. You can import other people’s themes by simply copying their Python script into your Blender/scripts directory and restarting Blender. Once Blender has restarted, switch to the Scripts window, click Scripts » Themes and select the theme you want. Close Blender. When you restart Blender, you will see your themes listed in User Preferences window, Themes tab. Finally, press CtrlU to save it as your default.
To preserve your current theme when opening a new file, disable the Load UI button.
Default
All images in this wiki have been screenshot using the Default icons and theme, so that it all looks consistent and less confusing. If you are looking at a private tutorial, or a forum sceenshot, keep in mind they might have changed all the colors and icons, and you will have to match up based on control placement and name.
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Customizing Theme
Click on a custom theme from the selection box, or Add a copy of the default one. When you do, many more columns of buttons will be shown. In the second column, you have in order:
- The name of the theme. Click into this field to edit it.
- The section of the theme to change, mostly organized by window type.
- The element within that section to customize (e.g. the background or text color…).
The next column over shows the current settings for that element (often color, but also sometimes size, etc.).
You can export your edited theme as a Python file using the File » Export » Save Current Theme... menu entry (which by the way calls a python script…).